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Int J Mol Sci 22:8126 Gradogna A, Lagostena L, Beltrami S, Tosato E, Picco C, Scholz-Starke J, Sparla F, Trost P, Caraneto A (2023) Tonoplast cytochrome b 561 is a transmembrane ascorbate-dependent monodehydroascorbate reductase: functional characterization of electron currents in plant vacuoles